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Unified Medical Language System® (UMLS®)

2012AB Gene Ontology Source Information


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Attribute Name Description Count (MRSAT.RRF)
GXR GO Cross Reference to external databases (e.g. MetaCyc:TRNA-CHARGING-PWY) 45845
GO_NAMESPACE Go Namespace field data 36343
REF References or citations related to a given atom or concept. 33963
GO_SUBSET Go Subset field data 9622
DATE_CREATED Date created 7860
GO_COMMENT GO Comment field data 3839
SID Secondary GO ID (e.g. GO:0020034) 1565
SYN_QUALIFIER Synonym qualifier 72
DISJOINT_FROM Disjoint from 6


  • Attributes included every discrete piece of information about a concept, an atom, or a relationship that is not part of the basic Metathesaurus concept structure or distributed in one of the relationship files.
  • Attribute Names (ATN) are based on source documentation or NLMs understanding of the source.
  • Sample attribute values (ATV) are provided for each ATN in the source.
  • Sample data highlight examples of ATNs and Attribute Values (ATV) in MRSAT.RRF.
  • See Metathesaurus Documentation for a complete list of attribute names.
  • See Section 2.5 of the UMLS Reference Manual for more information on attributes in the UMLS Metathesaurus.

Sample Data


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C1150869 A11607145 L2429914 S2867281 SDUI GO:0004421 GXR EC: hydroxymethylglutaryl-CoA synthase activity
C1152455 A11601883 L2427443 S2864552 SDUI GO:0008444 GXR EC: CDP-diacylglycerol-glycerol-3-phosphate 3-phosphatidyltransferase activity
C1150788 A11620367 L3493879 S4026396 SDUI GO:0008674 GXR EC: 2-dehydro-3-deoxy-6-phosphogalactonate aldolase activity
C1151366 A11589900 L2431549 S2867535 SDUI GO:0015043 GXR MetaCyc:LEGHEMOGLOBIN-REDUCTASE-RXN leghemoglobin reductase activity
C1151704 A11621382 L3493854 S4026704 SDUI GO:0018509 GXR MetaCyc: cis-2,3-dihydrobiphenyl-2,3-diol dehydrogenase activity
C1150655 A11590436 L2428024 S2865275 SDUI GO:0018524 GXR EC:6.4.1.- acetophenone carboxylase activity
C1149703 A11627706 L2430244 S2867090 SDUI GO:0018786 GXR UM-BBD_enzymeID:e0003 haloalkane dehalogenase activity
C1156312 A11603012 L6346731 S7272235 SDUI GO:0018902 GXR UM-BBD_pathwayID:dcp 1,3-dichloro-2-propanol metabolic process
C1152538 A11585693 L2427301 S2866634 SDUI GO:0042557 GXR EC:2.7.1.- eukaryotic elongation factor-2 kinase activator activity
C1152364 A11604842 L2431369 S2867320 SDUI GO:0045140 GXR EC:2.4.1.- inositol phosphoceramide synthase activity
C1327367 A11630162 L3499743 S4022756 SDUI GO:0046897 GXR RESID:AA0338 N-terminal peptidyl-tyrosine methylation
C1324336 A11611659 L3494253 S4026253 SDUI GO:0047090 GXR EC: benzoyl-CoA 3-monooxygenase activity
C1324649 A11611675 L3493909 S4024332 SDUI GO:0047189 GXR MetaCyc: 2,3-diaminopropionate N-oxalyltransferase activity
C1325120 A11617745 L3494503 S4024768 SDUI GO:0047345 GXR MetaCyc: ribose-5-phosphate adenylyltransferase activity
C1324257 A11611799 L3495970 S4028657 SDUI GO:0047985 GXR EC: hydrogen dehydrogenase activity
C1325207 A11605834 L3496494 S4022894 SDUI GO:0050154 GXR MetaCyc:OPHELINE-KINASE-RXN opheline kinase activity
C1324749 A11612014 L3495527 S4022921 SDUI GO:0050156 GXR EC: ornithine N-benzoyltransferase activity
C1323570 A11587140 L3495798 S4026895 SDUI GO:0050525 GXR EC: cutinase activity
C1150605 A11614236 L2431455 S2864823 SDUI GO:0008545 GXR EC:2.7.11.- JUN kinase kinase activity


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C1755622 A11581684 L3499785 S4022779 SDUI GO:0000398 GO_NAMESPACE biological_process nuclear mRNA splicing, via spliceosome
C1820103 A11581688 L6344094 S7270135 SDUI GO:0000405 GO_NAMESPACE molecular_function bubble DNA binding
C1523336 A11588077 L5382676 S6156215 SDUI GO:0001876 GO_NAMESPACE molecular_function lipoarabinomannan binding
C1622926 A11625309 L5829151 S6663196 SDUI GO:0001940 GO_NAMESPACE cellular_component male pronucleus
C1622390 A11594395 L5820704 S6665996 SDUI GO:0001970 GO_NAMESPACE biological_process positive regulation of activation of membrane attack complex
C1820155 A11588299 L6347203 S7276470 SDUI GO:0002925 GO_NAMESPACE biological_process positive regulation of humoral immune response mediated by circulating immunoglobulin
C1149754 A11600858 L2429360 S2865779 SDUI GO:0003837 GO_NAMESPACE molecular_function beta-ureidopropionase activity
C1150018 A11600895 L2428540 S2865366 SDUI GO:0004013 GO_NAMESPACE molecular_function adenosylhomocysteinase activity
C1151968 A11625641 L2427499 S7270005 SDUI GO:0004315 GO_NAMESPACE molecular_function 3-oxoacyl-[acyl-carrier-protein] synthase activity
C1152904 A11588560 L2429116 S2864946 SDUI GO:0004973 GO_NAMESPACE molecular_function N-methyl-D-aspartate receptor-associated protein activity
C1156526 A11595102 L6346726 S7272182 SDUI GO:0006072 GO_NAMESPACE biological_process glycerol-3-phosphate metabolic process
C1326498 A11595492 L3498044 S4027494 SDUI GO:0007395 GO_NAMESPACE biological_process dorsal closure, spreading of leading edge cells
C1151833 A11608121 L2429122 S2864579 SDUI GO:0008445 GO_NAMESPACE molecular_function D-aspartate oxidase activity
C1152684 A11626609 L2430631 S2866552 SDUI GO:0008529 GO_NAMESPACE molecular_function endogenous peptide receptor activity
C1151223 A11626683 L2430282 S2866880 SDUI GO:0008875 GO_NAMESPACE molecular_function gluconate dehydrogenase activity
C1157221 A11602051 L6342935 S7270741 SDUI GO:0009108 GO_NAMESPACE biological_process coenzyme biosynthetic process
C1157363 A11614432 L6343092 S7272225 SDUI GO:0009247 GO_NAMESPACE biological_process glycolipid biosynthetic process
C1325406 A11595913 L3494133 S4025506 SDUI GO:0009495 GO_NAMESPACE molecular_function thioredoxin-like 2Fe-2S ferredoxin
C1166831 A11608797 L2323037 S2745499 SDUI GO:0016011 GO_NAMESPACE cellular_component dystroglycan complex


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C2256913 A14235000 L7600147 S8796138 AUI GO:0003869 REF EC: PNPPase activity
C2257889 A14250813 L7605540 S8810007 AUI GO:0004069 REF EC: glutamic--oxaloacetic transaminase activity
C2259969 A14266821 L7598619 S8816397 AUI GO:0004347 REF EC: oxoisomerase activity
C2260009 A14240530 L7609302 S8809992 AUI GO:0004352 REF EC: glutamic acid dehydrogenase
C2261507 A14256568 L7622442 S8823793 AUI GO:0004674 REF EC: twitchin kinase activity
C2262104 A14272547 L7620859 S8810388 AUI GO:0004820 REF EC: glycyl translase activity
C1151271 A14256774 L7615406 S8795408 AUI GO:0004854 REF EC: NAD-xanthine dehydrogenase activity
C2263591 A14262241 L7608700 S8803615 AUI GO:0008444 REF EC: cytidine 5'-diphospho-1,2-diacyl-sn-glycerol (CDPdiglyceride):sn-glycerol-3-phosphate phosphatidyltransferase activity
C2263854 A14246450 L7606840 S8821444 AUI GO:0008609 REF EC: DHAP-AT
C2264132 A14251762 L7610508 S8820753 AUI GO:0008742 REF EC: ribulose phosphate 4-epimerase activity
C2264160 A14267564 L7611527 S8795713 AUI GO:0008747 REF EC: NANA lyase activity
C2264328 A14246525 L7610761 S8822735 AUI GO:0008791 REF EC: succinyl-CoA:L-arginine 2-N-succinyltransferase activity
C2261767 A14241402 L2432221 S2868956 AUI GO:0008998 REF EC: ribonucleotide reductase activity
C2265304 A14273121 L2326604 S2748858 AUI GO:0009703 REF EC: nitrate reductase (NADH2)
C2244820 A14258005 L7596384 S8796418 AUI GO:0030785 REF EC: RuBisCO LSMT activity
C1150005 A14242849 L7625840 S8820788 AUI GO:0033885 REF EC: sEH
C2247471 A14253392 L7625685 S8819469 AUI GO:0033951 REF EC: reducing end xylose-releasing exo-oligoxylanase activity
C2247630 A14253456 L7602938 S8820661 AUI GO:0034062 REF EC: ribonucleate nucleotidyltransferase activity
C1752425 A14258990 L5382837 S6157500 AUI GO:0040014 REF GOC:dph, GOC:tb regulation of body growth


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C1817202 A11588080 L6345378 S7270943 SDUI GO:0001897 GO_SUBSET gosubset_prok cytolysis by symbiont of host cells
C1148693 A11613235 L2429046 S2864632 SDUI GO:0003906 GO_SUBSET gosubset_prok DNA-(apurinic or apyrimidinic site) lyase activity
C1150497 A11619326 L2431480 S2864972 SDUI GO:0003951 GO_SUBSET gosubset_prok NAD+ kinase activity
C1152484 A11600918 L3496525 S4021722 SDUI GO:0004125 GO_SUBSET gosubset_prok L-seryl-tRNASec selenium transferase activity
C1152039 A11619396 L2431019 S2866965 SDUI GO:0004364 GO_SUBSET gosubset_prok glutathione transferase activity
C1150678 A11625705 L2430109 S6665580 SDUI GO:0004632 GO_SUBSET gosubset_prok phosphopantothenate--cysteine ligase activity
C1151789 A11582267 L2430274 S4024869 SDUI GO:0004755 GO_SUBSET gosubset_prok saccharopine dehydrogenase (NADP+, L-glutamate-forming) activity
C1156467 A11595065 L6351885 S7273501 SDUI GO:0006013 GO_SUBSET gosubset_prok mannose metabolic process
C1157109 A11595126 L6349835 S7277050 SDUI GO:0006145 GO_SUBSET gosubset_prok purine base catabolic process
C0683142 A11619850 L6348353 S7275405 SDUI GO:0006259 GO_SUBSET goslim_plant DNA metabolic process
C1817270 A11588931 L6350062 S7278986 SDUI GO:0006313 GO_SUBSET gosubset_prok transposition, DNA-mediated
C1157859 A11620013 L6345510 S7272154 SDUI GO:0006751 GO_SUBSET gosubset_prok glutathione catabolic process
C1157257 A11620036 L6350548 S7272365 SDUI GO:0006785 GO_SUBSET gosubset_prok heme b biosynthetic process
C1150930 A11583208 L2427264 S2864262 SDUI GO:0008684 GO_SUBSET gosubset_prok 2-oxopent-4-enoate hydratase activity
C1150044 A11589455 L2429690 S2866028 SDUI GO:0008810 GO_SUBSET gosubset_prok cellulase activity
C1150799 A11602009 L2428708 S2869347 SDUI GO:0009025 GO_SUBSET gosubset_prok tagatose-bisphosphate aldolase activity
C1158580 A11595776 L6348330 S7275244 SDUI GO:0009116 GO_SUBSET gosubset_prok nucleoside metabolic process
C1158633 A11595812 L6347848 S7271117 SDUI GO:0009200 GO_SUBSET gosubset_prok deoxyribonucleoside triphosphate metabolic process
C1157064 A11620660 L6346478 S7270017 SDUI GO:0009708 GO_SUBSET gosubset_prok benzyl isoquinoline alkaloid biosynthetic process


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C3158293 A19351890 L9977204 S12466713 SDUI GO:0001067 DATE_CREATED 2010-10-21T04:08:56Z regulatory region nucleic acid binding
C2753525 A17476000 L9161289 S11416584 SDUI GO:0003127 DATE_CREATED 2009-09-22T02:45:48Z detection of nodal flow
C2753690 A17472745 L9161157 S11415471 SDUI GO:0003294 DATE_CREATED 2009-10-22T08:35:36Z atrial ventricular junction remodeling
C2756017 A17472894 L6343986 S7280260 SDUI GO:0023058 DATE_CREATED 2010-02-16T09:30:50Z adaptation of signaling pathway
C2752233 A17475190 L9161445 S11418016 SDUI GO:0035322 DATE_CREATED 2010-02-24T02:05:14Z mesenchymal cell migration involved in limb bud formation
C2752335 A17472996 L9158073 S11418548 SDUI GO:0035417 DATE_CREATED 2010-03-29T10:53:42Z negative regulation of mitotic prometaphase
C3158746 A19349534 L9976508 S12459941 SDUI GO:0035572 DATE_CREATED 2010-08-06T02:00:17Z N-terminal peptidyl-serine dimethylation
C3158881 A19344155 L9978596 S12462837 SDUI GO:0035694 DATE_CREATED 2011-02-28T11:06:40Z mitochondrial protein catabolic process
C3159076 A19351004 L9974947 S12462410 SDUI GO:0035873 DATE_CREATED 2011-05-31T09:26:59Z lactate transmembrane transport
C2752547 A17477514 L9155110 S11415503 SDUI GO:0044295 DATE_CREATED 2010-02-04T03:57:16Z axonal growth cone
C3159211 A19342864 L9978398 S12460997 SDUI GO:0044331 DATE_CREATED 2010-08-20T10:24:54Z cell-cell adhesion mediated by cadherin
C2752780 A17477605 L9156435 S11419469 SDUI GO:0060557 DATE_CREATED 2009-04-13T02:46:14Z positive regulation of vitamin D biosynthetic process
C2752786 A17470949 L9155461 S11418700 SDUI GO:0060563 DATE_CREATED 2009-04-29T01:50:05Z neuroepithelial cell differentiation
C2752985 A17474293 L9161134 S11415201 SDUI GO:0060757 DATE_CREATED 2009-06-22T02:50:56Z adult foraging behavior
C2753001 A17470982 L9155285 S11417011 SDUI GO:0060773 DATE_CREATED 2009-07-25T06:28:38Z flower phyllotactic patterning
C2753014 A17472117 L9161662 S11419891 SDUI GO:0060786 DATE_CREATED 2009-07-31T01:55:19Z regulation of cell differentiation involved in tissue homeostasis
C2753029 A17473211 L9155455 S11418657 SDUI GO:0060801 DATE_CREATED 2009-08-06T01:24:24Z negative regulation of trophoblast cell differentiation by transforming growth factor beta signaling pathway
C2753072 A17477655 L9156866 S11415437 SDUI GO:0060844 DATE_CREATED 2009-08-12T10:49:00Z arterial endothelial cell fate commitment
C2753077 A17476539 L9155640 S11420285 SDUI GO:0060849 DATE_CREATED 2009-08-12T11:19:14Z regulation of transcription involved in lymphatic endothelial cell fate commitment


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C1151041 A11625087 L2427458 S2864525 SDUI GO:0000252 GO_COMMENT Note that zymosterol is cholesta-8,24-dien-3-ol. C-3 sterol dehydrogenase (C-4 sterol decarboxylase) activity
C1327242 A11625118 L3496962 S4024473 SDUI GO:0000380 GO_COMMENT Note that this process most commonly occurs in cis, selecting or skipping exons from the same primary transcript, but it has also been observed to occur in trans at low frequency, at least in some mammals. alternative nuclear mRNA splicing, via spliceosome
C1325278 A11607338 L0696000 S1205773 SDUI GO:0005208 GO_COMMENT This term was made obsolete because it represents a gene product. amyloid protein
C1151148 A11621182 L2428296 S4022965 SDUI GO:0016614 GO_COMMENT Note that enzymes of class EC:1.1.99.- should also be annotated to this term. oxidoreductase activity, acting on CH-OH group of donors
C1148668 A11627516 L2430897 S2864697 SDUI GO:0016917 GO_COMMENT See also the molecular function term 'neurotransmitter receptor activity ; GO:0030594'. GABA receptor activity
C1818047 A11590867 L6351326 S7279201 SDUI GO:0021521 GO_COMMENT This term was added by GO_REF:0000021. ventral spinal cord interneuron specification
C1818067 A11609558 L6351584 S7270796 SDUI GO:0021540 GO_COMMENT This term was added by GO_REF:0000021. corpus callosum morphogenesis
C1818106 A11609571 L6351201 S7278183 SDUI GO:0021573 GO_COMMENT This term was added by GO_REF:0000021. rhombomere 7 development
C1818124 A11584607 L6349411 S7273553 SDUI GO:0021582 GO_COMMENT This term was added by GO_REF:0000021. medulla oblongata maturation
C1818179 A11590894 L6343440 S7275311 SDUI GO:0021628 GO_COMMENT This term was added by GO_REF:0000021. olfactory nerve formation
C1818307 A11584632 L6352464 S7278544 SDUI GO:0021741 GO_COMMENT This term was added by GO_REF:0000021. spinal trigeminal nucleus development
C1818313 A11584633 L6347802 S7270715 SDUI GO:0021747 GO_COMMENT This term was added by GO_REF:0000021. cochlear nucleus development
C1818345 A11597206 L6352081 S7275316 SDUI GO:0021779 GO_COMMENT This term was added by GO_REF:0000021. oligodendrocyte cell fate commitment
C1818349 A11584636 L6349813 S7276797 SDUI GO:0021783 GO_COMMENT This term was added by GO_REF:0000021. preganglionic parasympathetic nervous system development
C1818400 A11615714 L6347755 S7270397 SDUI GO:0021832 GO_COMMENT This term was added by GO_REF:0000021. cell-cell adhesion involved in cerebral cortex tangential migration using cell-cell interactions
C1818408 A11603452 L6350423 S7271296 SDUI GO:0021840 GO_COMMENT This term was added by GO_REF:0000021. directional guidance of interneurons involved in migration from the subpallium to the cortex
C1818425 A11621882 L6350570 S7272570 SDUI GO:0021856 GO_COMMENT This term was added by GO_REF:0000021. hypothalamic tangential migration using cell-axon interactions
C1818453 A11597218 L6349034 S7270324 SDUI GO:0021883 GO_COMMENT This term was added by GO_REF:0000021. cell cycle arrest of committed forebrain neuronal progenitor cell
C1818484 A11590954 L6349951 S7278035 SDUI GO:0021912 GO_COMMENT This term was added by GO_REF:0000021. regulation of transcription from RNA polymerase II promoter involved in spinal cord motor neuron fate specification


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C1156150 A11587844 L5827806 S6668546 SDUI GO:0000055 SID GO:0000057 ribosomal large subunit export from nucleus
C1160083 A11625274 L5384080 S6156702 SDUI GO:0001841 SID GO:0001679 neural tube formation
C1817384 A11594428 L6346610 S7271059 SDUI GO:0002213 SID GO:0002214 defense response to insect
C1148886 A11600841 L2320539 S2752094 SDUI GO:0003727 SID GO:0003728 single-stranded RNA binding
C1149552 A11582071 L2429777 S2866048 SDUI GO:0003754 SID GO:0003758 chaperone activity
C1151515 A11600909 L2429636 S2865991 SDUI GO:0004096 SID GO:0016952 catalase activity
C1149911 A11607147 L2427591 S2868438 SDUI GO:0004438 SID GO:0016315 phosphatidylinositol-3-phosphatase activity
C1149327 A11601095 L2430366 S2867643 SDUI GO:0005041 SID GO:0008032 low-density lipoprotein receptor activity
C1817231 A11588644 L2428187 S2867983 SDUI GO:0005295 SID GO:0005282 neutral amino acid:sodium symporter activity
C1166607 A11625888 L2322496 S2744499 SDUI GO:0005575 SID GO:0008372 cellular_component
C1155661 A11582682 L2326483 S2748164 SDUI GO:0006298 SID GO:0006300 mismatch repair
C0597295 A11582707 L0040712 S1240511 SDUI GO:0006412 SID GO:0006416 translation
C1519613 A11601476 L2325597 S2752932 SDUI GO:0006413 SID GO:0006440 translational initiation
C1157562 A11582711 L2324378 S2752930 SDUI GO:0006414 SID GO:0006455 translational elongation
C0596562 A11607748 L6344271 S7271677 SDUI GO:0006633 SID GO:0000037 fatty acid biosynthetic process
C1155722 A11582914 L5825627 S6663659 SDUI GO:0007079 SID GO:0007082 mitotic chromosome movement towards spindle pole
C1151628 A11601961 L2429558 S4026437 SDUI GO:0008805 SID GO:0047767 carbon-monoxide oxygenase activity
C1158615 A11595852 L6347850 S7271120 SDUI GO:0009262 SID GO:0009122 deoxyribonucleotide metabolic process
C1159339 A11618772 L1218132 S2750816 SDUI GO:0009306 SID GO:0045166 protein secretion


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C2266398 A14250355 L7619975 S8801785 AUI GO:0001525 SYN_QUALIFIER systematic synonym blood vessel formation from pre-existing blood vessels
C2244203 A14271499 L7620727 S8809234 AUI GO:0001756 SYN_QUALIFIER systematic synonym formation of mesodermal clusters
C2246013 A14245128 L7617436 S8813540 AUI GO:0001946 SYN_QUALIFIER systematic synonym lymph vessel formation
C2263200 A14272722 L7609266 S8809644 AUI GO:0007283 SYN_QUALIFIER systematic synonym generation of spermatozoa
C2263202 A14267391 L7607475 S8827849 AUI GO:0007296 SYN_QUALIFIER systematic synonym yolk production
C2263225 A14272734 L7598540 S8815696 AUI GO:0007409 SYN_QUALIFIER systematic synonym neuron long process generation
C2265241 A14267750 L5827202 S6663531 AUI GO:0009556 SYN_QUALIFIER systematic synonym microspore development
C1160491 A14257340 L7605345 S8808213 AUI GO:0009561 SYN_QUALIFIER systematic synonym embryo sac development from the megaspore
C2265292 A14241479 L7606402 S8817330 AUI GO:0009640 SYN_QUALIFIER systematic synonym plant development in response to light
C2244306 A14252534 L7625257 S8815670 AUI GO:0022008 SYN_QUALIFIER systematic synonym nervous system cell generation
C2248107 A14264126 L7597818 S8809769 AUI GO:0042063 SYN_QUALIFIER systematic synonym glial cell generation
C2253860 A14254784 L7610012 S8816310 AUI GO:0048477 SYN_QUALIFIER systematic synonym ovum development
C2253996 A14265349 L7609858 S8815032 AUI GO:0048769 SYN_QUALIFIER systematic synonym myofibril production
C2262715 A14271344 L7613992 S8817461 AUI GO:0055046 SYN_QUALIFIER systematic synonym pollen development from the microspore
C0030012 A14271357 L7625339 S8816384 AUI GO:0055114 SYN_QUALIFIER systematic synonym oxidoreductase process
C2611006 A16504577 L8612419 S10714603 AUI GO:0070309 SYN_QUALIFIER systematic synonym lens fiber cell morphogenesis during differentiation
C2611164 A16502472 L8612752 S10717713 AUI GO:0070413 SYN_QUALIFIER systematic synonym trehalose metabolic process involved in response to stress
C2611165 A16502474 L8617652 S10717712 AUI GO:0070414 SYN_QUALIFIER systematic synonym trehalose metabolic process involved in response to heat stress
C2611166 A16502475 L8614837 S10717711 AUI GO:0070415 SYN_QUALIFIER systematic synonym trehalose metabolic process involved in response to cold stress


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C1148560 A11607042 L2324874 S2748253 SDUI GO:0003674 DISJOINT_FROM GO:0005575 ! cellular_component molecular_function
C1148560 A11607042 L2324874 S2748253 SDUI GO:0003674 DISJOINT_FROM GO:0008150 ! biological_process molecular_function
C1166607 A11625888 L2322496 S2744499 SDUI GO:0005575 DISJOINT_FROM GO:0008150 ! biological_process cellular_component
C1325880 A11627008 L3498239 S4026547 SDUI GO:0009987 DISJOINT_FROM GO:0032501 ! multicellular organismal process cellular process
C1325880 A11627008 L3498239 S4026547 SDUI GO:0009987 DISJOINT_FROM GO:0051704 ! multi-organism process cellular process
C1327400 A11591625 L6351947 S7274082 SDUI GO:0032501 DISJOINT_FROM GO:0051704 ! multi-organism process multicellular organismal process